Increasingly in international research and popular media a growing interest in men and fatherhood is discernible. These changes occur as other aspects of the socio-economic world shift, necessitating the need to re-address how caring and paid work responsibilities are configured and practised. However, interest in men’s experiences as fathers has emerged in ways which reflect cultural assumptions and practices associated with dominant understandings of masculinities. Consequently, research on and evidence of changing behaviours has been culturally and geographically uneven. In this paper, two qualitative studies are drawn upon to examine how men living in Australia and the UK engage in/narrate experiences of preparation for first-time fatherhood. These studies compare men’s in-depth accounts of preparing for first-time fatherhood in cultures where understandings of masculinities overlap, but where differences are also discernible. The findings illuminate the ways in which biology, gender, temporality and histories of masculinities frame men’s preparation activities and service provision.